well the main thing is pH. micro nutes are locked out easily by a pH issue.
one thing is for sure, its a micro nutrient. Ive got my money on zinc or magnesium defiency. id lean to the zinc deficiency becuase they lack a shine to them. but the dead spots makes me lean towards mag def. also mag deficiency is common when pH is too low (acidic), which you are. you said 5.5-6.5. you need to stay closer to 6.5. i know 1pH doesnt sound like much, but 5.5 is 10 times more acidic then 6.5
could also be iron def in the early stages, but its better to add a little iron now, before it gets worse (if thats the case)
either way, its a micro nutrient. check your fertilizers, see if these 3 elements are listed. if one (or more) isnt listed on the bottle, then thats probably what your deficient in.
I bought a bottle at a pet store that was for aquatic plants, for a fish tank, had 0-0-0 NPK rating, but 14 micro nutes. FOURTEEN! i use it every feeding, and have never had any issues, from start to finish. paid like 8 bucks i think.
at the same time, i also use 4 different veg+bloom nutes from start to finish, at different ratios depending the stage of the plant, and each brings its own set of micro nutes.
youll be ok, just fix the pH, and slowly add in some micronutes. slowly, cuz if it was pH lockout, the nutes are there, just not able to be absorbed, you dont want an abundance.
**if you do go to the petstore to get some of that micro nutes bottle for plants that kicks so much ass, get some pH buffer, most are 7.0 but if your lucky, they wil have a bottle of 6.5 (usually for african cichlids) but EXCELLENT for pot in soil. i use that every feeding too, and i know that liquid going in is 6.5, and runoff is around 6.2ish)**