from your own citation, the only part which pertains to GMO's at all.
"A connection between Bt maize and CCD was raised in experiments conducted in Germany that were described on the Internet but never published in a scientific journal. In these studies honeybees were fed Bt maize pollen and, although healthy bees had no acute or chronic toxic symptoms, in one experiment where bees were infested with parasites, the study was aborted because Bt pollen appeared to accelerate the bees decline. Although not repeatable in subsequent experiments, Bt in GE corn pollen thus became a possible cause of CCD.[SUP][138][/SUP]
However, there are no data in the scientific literature supporting direct or indirect damage to bees caused by currently approved GE crops engineered to make Bt proteins. For example, in 2008 a meta-analysis[SUP][139][/SUP] of 25 independent studies assessing effects of Bt Cry proteins on honeybee survival (mortality) showed that Bt proteins used in commercialized GE crops to control lepidopteran and coleopteran pests do not negatively impact the survival of honeybee larvae or adults. Additionally, larvae consume only a small percent of their protein from pollen, and there is also a lack of geographic correlation between GE crop locations and regions where CCD occurs.[SUP][138][/SUP] "
only captive bees FED pollen could establish the link between GMO's and bees.
bees however do not eat corn pollen. bees visit flowers to collect nectar, and the pollen adheres to their bodies to be deposited on the next flower they visit, and random mixed pollen is stored in the hive to be converted into a fungal mass for protein. bees have no interest in corn pollen. even their pollen collection is a secondary action behind the collection of nectar for honey production.
the bees had to be FED corn pollen since in the wild bees do not interact with corn. corn pollinates via large (rice sized and larger) pollen packets which fertilize the leaf internodes to develop ears of grain. corn tassels do not provide nectar, and thus lack the primary attractant to make bees want to bother with corn. corn pollen is too large to adhere to bees, the packets are too large to be carried home by bees, and bees have less interest in corn than they do in marble statuary of kids pissing into fountains. the entire "GMO corn = bee colony collapse syndrome" hypothesis is ridiculous on it's face.
publishing this kind of twaddle gets journals laughed into obscurity, this is why it was never actually published, just shopped to eco-websites where dipshits who dont know how corn grows will accept this story as fact and repeat it to their gullible pals at "occupy" rallies.